Vitaceae

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Keys in Flora of Bhutan:
1. Leaves simple (but often lobed) ......... .. 2
+ Leaves compound ............................... .. 5
2. Tendrils usually twice bifid, never bome on peduncles; flowers 4-merous; style elongate ........... .. 5. Cissus
+ Tendrils once-bifid, or with one branch again bifid, often bome on peduncles; flowers 5-merous; style very shon or absent ........ .. 3
3. Flowers in rounded cymes: petals free and spreading at anthesis ..........  2. Ampelocissus
+ Flowers in panicles; petals free and spreading or united and shed as a cap at anthesis ......... .. 4
4. Stems, petioles and peduncles never softly glandular-bristly; petals united at apex and shed as a cap at anthesis ...... 1. Vitis
+ Stems, petioles and peduncles softly glandular-bristly; petals free at apex and spreading at anthesis ......................... .. 2. Ampelocissus (A. barbata)
5. Leaves trifoliolate ................................................ .. 6
+ Leaves palmately or pedately 4 — 7-foliolate ...... .. 9
6. Young stems whitish woolly; lateral leaflets often lobed; peduncles usually bearing a tendril; flowers 5-merous ........ .. 2. Ampelocissus (A. divaricatus)
+ Stems glabrous or pubescent but never whitish woolly; lateral leaflets unlobed; peduncles without tendril; flowers 4- or 5-merous ................ 7
7. Tendrils 2 - 3 x bifid, branches ending in disc-like pads; flowers in terminal and leaf-opposed cymes, 5-merous; style stout, shonly columnar  ............. 3. Parthenocissus
+ Tendrils simple or 1 — 2 x bifid, but branches not ending in disc-like pads; flowers in axillary or pseudo-axillary cymes, 4-merous; style absent or elongate and slender .................... .. 8
8. Tendrils simple; leaves glabrous beneath; stigma conspicuously 4-lobed or disc-like, sessile on ovary ............ .. 4. Tetrastigma
+ Tendrils 1 — 2 x bifid; leaves softly villous beneath; stigma minute on slender elongate style ............... .. 7. Cayratia (C. geniculata)
9. Leaves palmately compound; berry 1 - 2-seeded ................ 10
+ Leaves pedately compound; berry 2 — 4-seeded ............. .. 12
10. Tendrils simple or radiately branched; style absent or very shon ....... 4. Tetrastigma
+ Tendrils 1 — 2 x bifid; style elongate .................................................. .. 11
1 1. Young stems, petioles, inflorescences and leaves glabrous or almost so; leaflet margins sharply serrulate; cymes leaf-opposed; berry 2 — 2.5cm; seed c 16 x 10mm ............... .. 5.Cissus (C. elongata)
+ Young stems, petioles, inflorescences and leaves (beneath) softly pubescent; leaflet margins crenate; cymes pseudo-axillary or temtinal; berry c l.5cm diameter; seed c 8 x 6mm ............... .. 6. Cyphostemma
12. Tendrils simple or bifid; flowers functionally unisexual; stigma broadly 4- lobed on shon thick style or subsessile ...... .. 4. Tetrastigma
+ Tendrils 1 - 2 x bifid; flowers bisexual; stigma minute on slender elongate style ....... 7. Cayratia (C. japonica)



 
Plant Taxonomy By O P. Sharma (1993)- Details