Scleromitrion verticillatum

Scleromitrion verticillatum (L.) R.J.Wang, J. Trop. Subtrop. Bot. 22: 440 (2014). (syn: Hedyotis crassifolia Blume, nom. illeg.; Hedyotis hispida Retz.; Hedyotis hispida Roth, nom. illeg.; Hedyotis verticillata (L.) Lam.; Hedyotis verticillata var. barlettii Fosberg & Sachet; Hedyotis verticillata var. scaberrima Hatus.; Hedyotis wallichii Walp.; Oldenlandia angustifolia Benth., nom. illeg.; Oldenlandia hispida (Retz.) Lam.; Oldenlandia verticillata L.; Scleromitrion crassifolium Miq.; Scleromitrion hispidum (Retz.) Korth.);

Trop. & Subtrop. Asia to WC. Pacific as per WCSP;

Assam; Bangladesh; Borneo; Caroline Is.; China South-Central; China Southeast; East Himalaya; Gilbert Is.; Hainan; India; Jawa; Kazan-retto; Malaya; Maluku; Myanmar; Nansei-shoto; Nepal; Nicobar Is.; Philippines; Sulawesi; Sumatera; Taiwan; Thailand; Vietnam as per Catalogue of Life;


Annual often robust, but very variable. Stems 10-30cm, terete or subterete below 4 angled above, hispid. Leaves opposite sessile; Lamina elliptic or linear lanceolate, 20-55 x 1.5-10mm, acute or acuminate, base shortly atenuate, uppersurface with conspicuous epidermal cells, scabrid lower surface paler, scabrid with prominent, hispid midrib; stipules with long often brownish often filiform, sparsely hispid appendages 3-6.5mm. Flowers in sessile axillary verticilasters; pedicels c 1mm in fruit.Calyx 3-4.5mm; Tubes and lobes subequal, densely hispid. Corolla shortly campanulate; tube very shortwithout a ring of hairs; filaments long anthers almost reaching tips of corolla lobes. Stigma slightly overlapping anthers shortly bifid.Capsule ovoid, c 2mm, hispid, loculicidal on crown only.
Fl. May-December
On sandy soil mostly in Terai.
(Attributions- A.C.J Grierson & D.G Long. Flora of Bhutan. Published by RBGE and RGoB. 1987 from Bhutan Biodiversity Portal)


https://08511630493324166816.googlegroups.com/attach/3ef18df6a8217123/3._Spermacoce_sp._-_Leaf_IMG_4209.jpg?part=0.3&view=1&vt=ANaJVrGgjuEo4TCkBADJa1ytOP3zsujJEJMehFqnwzFjJnwR9h-NGMC9i-XZc9T-69gG2L_-dvJ5yXxB5QZBTOqmriZRkYa4ZQIQgcw6J3LVeVs1mLjDbaM
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https://08511630493324166816.googlegroups.com/attach/d02130ae01020f4c/4.__Spermacoce_sp._-_Inflorescence_IMG_4207.jpg?part=0.4&view=1&vt=ANaJVrEspErtTZmysruvYxS33KB-xyHgw6Jr1EZlgve56B7yF5rTBv7JNC_ZGkJw7xZpZ2l_9f-e3xIddDPFlEq50RWLy5M_HGr1EqeM6NPU-9a3C8yQAs8
 
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Spermacoce remota Lam. (?) from Assam. : Attachments (6). 3 posts by 2 authors. 
Attached images may be Spermacoce remota Lam. (?). Please ID the plant.
Date :06.11.2013
Location: Kamrup district (Assam)
Family : Rubiaceae
Genus & species : Spermacoce remota Lam. (??)
Habitat: Grows wild on hilly slopes
Habit : Herb 
Flower : White in colour
Fruits : Not seen 
efi page on Spermacoce remota ? 

I think it should be Spermacoce pusilla Wall. as per comparative images at Spermacoce and as per keys at Flora of China
I mean here S.remota is correct.
Thanks, ...,
I do not think it is Spermacoce remota as per images and details & references herein.
But it may be not be Spermacoce pusilla Wall. either.
May be some other genus ?
After through search in Flora of China, Flora of Nepal, Efi site etc., I feel it is closer to Scleromitrion tenelliflorum (Blume) Korth. (syn: Hedyotis tenelliflora Blume) as per
Catalogue of Life  Flora of China  FOC illustration  GBIF (high resolution specimens)

On further checking,
I think it should be Scleromitrion verticillatum (L.) R.J.Wang (syn: Hedyotis verticillata (L.) Lam.) as per differences between closely related species at
GBIF (high resolution specimens) FOC illustration  Flora of China 

https://08511630493324166816.googlegroups.com/attach/589da5c47a64503/for%20id..%20(6).JPG?part=0.4&view=1&vt=ANaJVrH58Nxc_iktsaVd8IXiwwyjW_GwUtS7zE2tTvqwLUhKiaPbRA9c5D3K5UY1-KT8E-JbLzltuDuH542yEZBVX9cj06sJfGoB6hVZh9vv4AkRB3RS3io
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Dehradun-Mussoorie Tour 2013:: Rubiaceae Herb for id from Dehradun
 : Attachments (5). 5 posts by 3 authors.
This prostrate herb with stiff, glabrous, opposite leaves was shot from Malsi Deer Park, Mussoorie Road, Dehradun..
leaves with spinous apex, bearing clusters of minute white flowers in their axils....
I believe this belongs to Rubiaceae.. please help to id...
It may be Metabolos rugosus (= Hedyotis rugosa). Not sure though. 
After through search in Flora of China, Flora of Nepal, Efi site etc., I feel it is closer to Scleromitrion tenelliflorum (Blume) Korth. (syn: Hedyotis tenelliflora Blume) as per
Catalogue of Life  Flora of China  FOC illustration  GBIF (high resolution specimens)
Hedyotis rugosa (Blume) Korth. does not have any distribution in India.

On further checking,
I think it should be Scleromitrion verticillatum (L.) R.J.Wang (syn: Hedyotis verticillata (L.) Lam.) as per differences between closely related species at
GBIF (high resolution specimens) FOC illustration  Flora of China 
Thanks … for providing useful links, I do agree that it is quite difficult to conclude based on these pics only..
But I am equally surprised that the leaves do not fully match with any of the available pics or description, as they are markedly stiff as well as pointed at tip, a feature which should not have been missed, if it belongs to any of these taxa...!!


https://groups.google.com/group/indiantreepix/attach/5db68037d7a8c/DSCN9874%20-%20Copy.JPG?part=0.1&authuser=0&view=1
 
7 April 2019/NSD-1 Herb for ID : 6 posts by 4 authors. Attachments (1)- 1 Mb. 
Please ID this small herb...
Umariya, M.P. Sept 2017

its not in focus. i could not discern the features of the flowers. do you have a better focus photo?

Hedyotis cynantha  

To me looks different from images at Hedyotis cyanantha

Pl. check with other species at Hedyotis & Oldenlandia

I think it should be Scleromitrion verticillatum (L.) R.J.Wang (syn: Hedyotis verticillata (L.) Lam.) as per differences between closely related species at
GBIF (high resolution specimens) FOC illustration  Flora of China 

I guess it is different.
Pl. see variations at GBIF (high resolution specimens)
Scleromitrion tenelliflorum (Blume) Korth. ??
Calyx is densely hispidulous in Scleromitrion verticillatum (L.) R.J.Wang (syn: Hedyotis verticillata (L.) Lam.) and glabrous in Scleromitrion tenelliflorum (Blume) Korth. as per Scleromitrion sirayanum (Rubiaceae: Spermacoceae), a new species of the Hedyotis-Oldenlandia complex in Taiwan- Tian-Chuan HSU, and Zhi-Hao CHEN- Taiwania 62(2): 151‒156, 2017
So it is Scleromitrion verticillatum (L.) R.J.Wang (syn: Hedyotis verticillata (L.) Lam.) only.


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SK2080 27 July 2019 : 13 posts by 4 authors. Attachments (10)- around 600 kb each. 
Location:  Phedi, Nagarkot, Bhaktapur
Date: 6 July 2019
Elevation:  1647 m.
Habit : Wild
Does not seem any Spermacose (Borreria) listed in the ACFPN.

Some thing similar was identified while I was sorting out Spermacoce species under Spermacoce pusilla as Oldenlandia ?
However, I am not able to get hold of that post or species right now.

It resembles Oldenlandia nitida


Please see this old post of mine... I think they are similar.. but the id was not resolved

It looks like very much matching with your images but not sure about the ID,
and it is not listed in Nepal. How can we verify?
Scleromitrion verticillatum (L.) R.J.Wang
Syn : Hedyotis verticillata (L.) Lam.

After through search in Flora of China, Flora of Nepal, Efi site etc., I feel it is closer to Scleromitrion tenelliflorum (Blume) Korth. (syn: Hedyotis tenelliflora Blume) as per
Catalogue of Life  Flora of China  FOC illustration  GBIF (high resolution specimens)
Hedyotis rugosa (Blume) Korth. does not have any distribution in India.
It is quite like to be found in your area as WCSP shows distribution as Trop. & Subtrop. Asia to N. Queensland. Catalogue of Life gives it as Assam; Bangladesh; Borneo; China South-Central; China Southeast; East Himalaya; Hainan; Japan; Jawa; Korea; Malaya; Maluku; Myanmar; Nansei-shoto; New Guinea; Nicobar Is.; Philippines; Queensland; Sumatera; Taiwan; Thailand; Vietnam
Nidhan ji has posted the same plant from from Malsi Deer Park, Mussoorie Road, Dehradun
Here is a paper showing differences between confusingly similar species
Scleromitrion sirayanum (Rubiaceae: Spermacoceae), a new species of the Hedyotis-Oldenlandia complex in Taiwan- Tian-Chuan HSU, and Zhi-Hao CHEN- Taiwania 62(2): 151‒156, 2017- (ABSTRACT: The new species Scleromitrion sirayanum T.C. Hsu & Z.H. Chen (Rubiaceae) is described and illustrated from Taiwan. S. sirayanum is similar to S. angustifolium (Cham. & Schltdl.) Benth. but easily distinguished by its constantly solitary flowers, salverform corolla, adaxially basally long villous corolla lobes, spreading to ascending persistent calyx lobes, and obconic capsules dehiscent both septicidally and loculicidally into 4 valves. S. sirayanum is so far only found in Nanhua District of Tainan City and evaluated as Vulnerable based on the IUCN Red List criteria. Three new combinations are also proposed for the Hedyotis-Oldenlandia complex in Taiwan)
Scleromitrion angustifolium (Cham. & Schltdl.) Benth. is treated as a syn. of Scleromitrion tenelliflorum (Blume) Korth. in Catalogue of Life;
On further checking, I think it should be Scleromitrion verticillatum (L.) R.J.Wang (syn: Hedyotis verticillata (L.) Lam.) as per differences between closely related species at
GBIF (high resolution specimens) FOC illustration  Flora of China 

Thank you … for validation !



References:

Differences between closely related species at Scleromitrion sirayanum (Rubiaceae: Spermacoceae), a new species of the Hedyotis-Oldenlandia complex in Taiwan (hispidulous calyx for Scleromitrion verticillatum (L.) R.J.Wang)
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