Arisaema griffithii aff. griffithii-group

Arisaema griffithii aff. griffithii-group; 
 

Images by Ushaprabha Page & Pravin Kawale (Both Ids by Pascal Bruggeman) (Inserted by J.M.Garg) (For more images & complete details, click on the links) 

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id pl. of Arisema sp. : Attachments (6). 14 posts by 9 authors.
Which Arisaema-id pl.
Location- Eagle nest sanct. Altitude 9500feet.
time12.35 p.m.on 22nd April 10.

Cobra lily
Arisaema intermedium may be
I think ... is correct
I think this one should be Arisaema tortuosum.
Normally the whip-like cord coming from inside the throat (spadix appendage) is upright, but when the flower is past its prime, the cord droops, as seen in the pictures here.
It should not be Arisaema intermedium because in that species the "hood" (spathe) of the cobra-lily has a 2-3 cm long tail-like tip, as can be seen in the following:
http://www.pacificbulbsociety.org/pbswiki/index.php/ArisaemaSpeciesThree
The "hood" of Arisaema tortuosum, on the other hand, is pointed, but has no tail-like tip. To me, ... images appear to agree with Arisaema tortuosum.
..., the leaves are broadly trifoliolate, and not digitate as in case of A. tortuosum. I too have seen this plant (certainly not A. tortuosum) at the Eaglenest pass and wondering about its ID.
I can't make out the leaves clearly from the picture. If they are indeed trifoliate, it surely cannot be A. tortuosum.
Then, this could be a species not yet reported from Arunachal Pradesh.
We found Arisaema elephas in Tawang region, from where it is not reported (as far as I know). So, one may have to look at some species from China, the neighboring region.
One such candidate is Arisaema handelii, which has trifoliate leaves,
http://www.aroid.org/genera/arisaema/herold/Images/arhandelii.jpg
http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200027206
" Spathe green, purple, or dark green with broad white stripe", "Appendix whitish, narrowly conic, basal 4-6 cm densely papillose or rugose, ca. 1.5 cm in diam., abruptly narrowed at base into stipe ca. 6 × 5 mm, distal part smooth and gradually drawn into flagellum to 38 cm in total, exserted from spathe tube and tortuous, pendulous in distal flagellate part. "
Arisaema hookeri
May be A. sivdasani
This is a species that shows intermediate characteristics between the Arisaema griffithii-group (notice the reticulated innerside of spathe blade in the last picture) and the elephas alliance. AP is and area where the Sino-Himalayan and Indo-Himalayan floras meet and there are many plants that show characteristics of species from both areas. The Arisaema griffithii group usually has long threadlike appendices and not the thicker and short appendix of the plants on the pictures.
Similar but darker plants can be found in the Yumthang area of N Sikkim where the slow change from griffithii type plants to this type and further on to true Arisaema elephas is more apparant. Further east in AP similar plants like this are found that have a more beige-orange flowers.Some remarks on names mentioned in this thread:

- Arisaema handelli is very similar to Arisaema elephas but has a rather narrow distribution range in China, most important characteristic of this species is a rugose (rough, like sandpaper) appendix wheras the plants on the picture have a smooth appendix, it also has a flowerstalk that generally is shorter than the leafstalks
whereas the plant on the pictures has its flowers as high or higher than the leaves.
- Arisaema intermedium has a very long threadlike appendix that can reach 50-70 cm in length, it is also much smaller with narrow, more elliptic leaflets, not the broad rhombic leaflets of the AP plants
- Arisaema hookeri (or hookerianum) is the lowland form of Arisaema griffithii that can be found at Tiger Hill and some parts of Shingallila National Park in W Bengal. It differs from normal griffithii by a very triangular folded spathe and a more purple colour. Nevertheless it is a full synonym of Arisaema griffithii.
- Arisaema sivadasanii is a endemic autumn-flowering form of Arisaema tortuosum from S India. It has now been reduced to a variety of tortuosum by Punekar and Kumaran

Whether the plant on the pictures has characteristics sufficently unique to warrant a separate status extensive fieldwork needs to show.
As it is it doesn't fall within any known species I would suggest to ID it as Arisaema species aff. griffithii-group.


This Arisaema has been observed in various colour forms in AP but does not seem to fall in any of the know species. Because of the appendix shape it shows some resemblance with Arisaema elephas (which generally
is purple flowered) but particularly in the last picture the reticulation on the inside of the spathe blade shows its close affinity with members of the Arisaema griffithii group. Griffithii normally has a thin, threadlike appendix but in some areas at higher elevations the appendices seem to get shorter and thicker and slide
towards Arisaema elephas. This occurs for instance in the Yumthang area of N Sikkim. Because of the present reticulation (nettled veins) I would suggest to id it as Arisaema griffithii aff.
Further field research is needed to establish its true status and find out whether it deserves a separate status. Because Arunachal Pradesh is in between the Indo Himalayan and Sino Himalayan Flora many intermediate forms of Arisaema have been found of Indian and Chinese species, for instance intermediates between the sister species Arisaema galeatum (India) and Arisaema dahaiense (China).
Some remarks on the proposed names in this thread:
- As noted Arisaema handelii has a rugose (rough, like sandpaper) appendix and a rather restricted  distribution range in China, the plants on the pictures have a smooth appendix. A. handelii also has a
flower stalk shorter than the leaf stalks in contrast to the plants on the pictures.
- Arisaema sivadasanii is a narrow endemic species from S India and is recently reduced to a variety of Arisaema tortuosum, it does not occur in N India and is usually autumn flowering, it belong to a different
section within the genus and has pedatisect leaves, not tripartite
- Arisaema hookeri ("hookerianum) is a full synonym of Arisaema griffithii. It is the lowland form of griffithii from ~2000 m altitude that occurs for instance at Tiger Hill in W Bengal, it is usually reddish purple and of all the forms of griffithii has a spathe that is the most turned back on itself pointing much backward and usually even so recurved that it embrasses the flowering stalk.


 
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Arisaema costatum : Attachments (2).  6 posts by 5 authors.
Validate Please
Arisaema costatum
At Yumthang Valley,Sikkim
21May,2013
This is a narrow spathed variant of the griffithii alliance. With some variants it is difficult to assess whether they are still forms of griffithii with a narrow spathe or big forms of propinquum with a wider spathe as they seem part of a continuous variational range going from one extreme (griffithii) to the other (propinquum). I have studied this group in the Yumthang Valley and the variation is endless, particularly at the higher elevations and forms like this ultimately morph into elephas type plants. Due to this it is impossible to attach a definitive name to many plants from this valley other than that they belong to the “griffithii” group.
Costatum it is definitely not as that is a Nepal endemic with thick, shiny leaves with raised veins underneath growing at much lower altitudes.
Many people look at species as fixed entities but in reality several Arisaema are still in a high state of evolutionary flux which causes some problems with identifications. The best name for this plant would probably be Arisaema griffithii aff.
The same variational patterns for griffithii and allies occurs in the Shingallila National Park and several places in Nepal. 
For good photographs of Arisaema costatum in situ and the variation of the griffithii-group in Nepal see: http://www.greenmilenursery.be/photo_nepal.html

 
  
 
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