Kholia B. S.


List of Publications up to January 2015


P.S. Das  and B. S. Kholia 2014. A selective study of online resources of Information on plant taxonomy & systematics: A new path of data flow to the users in digital age. Indian Journal of Forestry 37(4) : 445-455.


B.S. Kholia and C. R. Fraser-Jenkins 2014.  Rediscovery of Ophioglossum pendulum L. from eastern Himalaya, with a note on historical background and new  distribution record to Sikkim. Indian Journal of Forestry 37(3): 347-354.



B. S. Kholia  2014. Cheilanthes tibetica  Fraser-Jenk. & Wangdi, A New fern to India. Indian J. For. 37: 179-182.



Durgesh Verma, S. K. Singh, B. S. Kholia, B. K. Sinha and Samiran Pandey 2014. Pteridophytes of Khawnglung wildlife sanctuary, Mizoram, India. Indian Journal of Forestry 37(2): 219-234.


B.S. Kholia, Piu Das and S. Pradhan 2014. On the occurrence of Athyrium nakanoi Makino (Woodsiaceae) in Sikkim Himalaya. Indian Journal of Forestry 37: 81-86.



B. S. KHOLIA 2014. Ferns and Fern-allies of Sikkim Himalaya –A Pictorial Handbook,  Part-II. Department of Forests, Environment, Wildlife management, Government of Sikkim and Botanical Survey of India Gangtok. pp. i-ix, 1-291.



B. S. Kholia. 2014. Traditional Uses of Pteridophytes in Sikkim Himalaya . in Sanjeev Kumar (ed) Ethanobotanical Studies in India  pp. 291-302. Deep Publications Delhi.


Srivastava, R., Uniyal,  P. L. and  B. S. Kholia 2014. Studies on Reproductive Biology of Microsorum alternifolium Copel. The International Journal of Plant Reproductive Biology 6(1) pp. 15-19.



Kholia, B. S. 2013. Sikkim Himalayan Tree Ferns: Diversity, Distribution, Uses, Economic Potentials And Conservation. In Ram Awtar Singh (Ed.). Geology, Biodiversity & Natural Resources of Himalaya and their intellectual property law. Gyanodaya Publication nainital. pp. 305-318. {ISBN: 978-93-82171-40-9}



B. S. Kholia . 2013. A new distribution record of filmy fern Hymenophyllum denticulatum Sw. from Uttarakhand Himalaya. Indian Fern J. 30 : 154-160 (2013).



B. S. KHOLIA. 2013 Occurrence of endangered orchid Cymbidium whiteae King and Pantl. in North Sikkim. Current Science 104 (1): 12-13,



B. S. KHOLIA. 2013.Note on the Re-collection and Conservation of Thelypteris kingii

(Thelypteridaceae) – An Endemic Fern of India. Taiwania, 58(1): 44–48, ISSN: 0372333X



Kholia, B.S., Ranjan Joshi  and Richa Punetha 2013. Extended distribution of Cyathea spinulosa Wall.ex Hook. in Uttarakhand Himalaya with a note on distribution and diversification of Himlayan ferns in relation to recent climate change. NeBIO. Vol. 4, ( 2) 40-45.



Punetha R, Bhakuni, K and Kholia, B.S. 2013. Lithophytic Ferns and Fern-allies of Kumaon. In (ed. R. A. Singh) Landslides and Environmental Degradation. Pp. 239-252. (ISBN: 81-85097-90-9). 



D. Verma, S. K. Singh, b. s. Kholia, B. K. Sinha and Samiran Panday 2013. Pteridophyte of Ngengpui Wildlife Sanctury, Mizoram India, Keanean Journal of Science 2: 3-12.



B. S. Kholia, Kamlesh Bhakuni, Richa Punetha and N. S. Bankoti 2012.Taxonomic studies on central Himalayan species of fern genus Lepisorus (Polypodiaceae) with a note on thickness of rhizome and deciduousness of the lamina. NeBIO. 3(3): 28-40.



Kamlesh Bhakuni, Richa Punetha and B. S. Kholia. 2012 On the reoccurrence of Pellaea calomelanos (sw.) Link, in central Himalaya with a note on distribution. (Ready for Nelumbo, The Bull. Bot. Surv. India 2013 (Just Published) ISSN, 0006-8128



B. S. KHOLIA. 2012.  Botrychium simplex E. Hitchcock: a New Moonwort for the Indian Himalayan Mountains. American Fern Journal 102(1):86-90. ISSN: 0002-8444.



B. S. KHOLIA  2012. Rediscovery of Trachycarpus martianus in Sikkim Himalaya. Palms the journal of International palm society. 56(1): 43-50. ISSN: 1523-4495



Joshi, R., Kumar, K. and Kholia, B.S. 2012. Adaptations for Tourism Management under Climate Change –Analyses of Sikkim Himalaya. NeBIO 3(4): 26-30.



B. S. KHOLIA and C. R. Fraser Jenkins 2011. Misidentification makes scientific publications worthless - Save our Taxonomy and Taxonomists. Current Science 100(4): 458-461.



B. S. KHOLIA 2011. Pteridophytic wealth of Sikkim Himalaya (Book Chapter) in Biodiversity of Sikkim ed.. M. L. Arrawatia, SandeepTambe and G. S. Rawat) Chapter 3.Ferns. 35-68. ISBN: 978-81-920437-0-9



B. S. KHOLIA and N. Punetha, 2011. Biology of ferns and Lycophytes  (Book review) Current Science 101(7): 963.



B. S. KHOLIA and N. Punetha, 2011. Fern Ecology (Book review) Current Science 100(12): 1885-1887.



B. S. KHOLIA, Kamalesh Bhakuni and R. Punetha 2010. Aleuritopteris punethae (Pteridaceae) a new species of silver fern from Indian Himalaya. Telopea 12 (4): 485-489. ISSN 0312-9764



B. S. KHOLIA 2010. Is Trachycarpus latisectus vanishing from its natural habitat. Palms The Journal of International Palm Society. 54(1): 48-55. ISSN: 1523-4495




B. S. KHOLIA 2010. Sikkim Silver brake at risk. Current Science 99(8): 999. ISSN 0011-3891



B. S. KHOLIA, Ranjan Joshi and U. Lachungpa 2010. International year of Biodiversity – Revisiting curricula for conservation Curr. Sci. 99(12):1637.



B. S. KHOLIA 2010. Kumaon fan palm: Trachycarpus takil Becc. (Arecaceae In Retrospect. ) (BOOK CHAPTE) In Y. P. S. Pangtey, LalitTiwari, and GeetaTiwari. Biodiversity potential in Himalaya. GyanodayaPrakashan Nainital. Pp.417-426.



B. S. KHOLIA 2010. On the correct identity of west Himalayan species of Plagiogyria  Nelumbo the Bull. Bot. Surv. India. 51: 223 – 224.



P. S. Dash and B. S. KHOLIA 2010. Indian Integrated Plant Taxonomic Information System: A conceptual frame work. DESIDOC. Journal of Library & Information Technology 30(3): 35-42.



B. S. KHOLIA and Ranjan Joshi 2010. International year of Biodiversity – Issues on some practices.  NEBio vol.2. pp. 56-61.



N. Punetha and B. S. KHOLIA 2010.  The Floristic Diversity of Uttarakhand: Ferns andFern-allies.  In The Plant Wealth of Uttarakhand.  Ed. Uniyal, P. L. B. P. Chamola & D. P. Semwal. 293-313. J. P. Publications New Delhi. (Book Chapter)



B. S. KHOLIA and Kamlesh Bhakumi 2009. Western Himalaya – a new range of distribution for a critically endangered fern Dryopsis manipurensis (Bedd.) Holttum & P.J. Edwards. Nelumbo the Bull. Bot. Surv. India. 51; 245 – 248.



B. S. KHOLIA. 2009. A note on mistaken identity of Sanjeevani Booti . Indian Fern J. Vol. 26: 199-200.



B. S. KHOLIA 2009. Gender variation in a threatened and endemic palm Trachycarpus takil  Becc. Current Science. 96(1): 144-148.



B. S. KHOLIA & A. A. Anshari  2009. On the occurrence of Aleuritopteris tambureii (Hook.) Ching in India. Nat. Acad. Sci. Lett. 32, NO.7 & 8, 22-230.



PUNETHA, N., KAMLESH BHAKUNI &  B. S. KHOLIA.2008.  On the occurrence of Cheilanthes argntea (a silver fern ) in Central Himalaya. J. Indian Bot. Soc. 87(3&4): 289-290. ISSN: 0019-4468



PUNETHA, N., R. PUNETHA & B. S. KHOLIA. 2008. Some aspects of morphology, branching ofrhizome and crozier uncoiling phenomenon in Osmunda regalis L. Phytomorphology. 58(1&2): 9-14. ISSN. 0031-9449



B. S. KHOLIA 2008.  Selaginella bryopteris (L) Bak. an Endemic Fern-ally of India under threat. Indian Fern J. 26: 73-78.



PUNETHA, N. , KAMLESH BHAKUNI & B. S. KHOLIA. 2008. Habitat ecology and Prothalial development in endemic Pteris subquinata . Indian J. For.  31(2): 287-290. (ISSN: 0971-9431



Gibbons, M. Spanner, Tobias and B. S. KHOLIA, 2008. Trachycarpus takil Becc. In Kumaon. Current Science. 94(4): 444-446.



B. S. KHOLIA & PUNETHA, N. 2007.On the taxonomic validity of Osmunda japonica Thunb. Indian Fern J. 24: 85-90. ISSN: 0970-2741



PUNETHA, N., KAMLESH BHAKUNI & B. S. KHOLIA. 2007. Insect fern interaction in central Himalaya. Nat. Acad. Sci. Lett. 30(9-10). ISSN: 0250-541X



B. S. KHOLIA & PUNETHA, N. 2005. Useful Pteridophytes of Kumaon. Indian Fern Journal. 1-6.



B. S. KHOLIA & PUNETHA, N. 2005Notes on Distribution and Conservation of the rare Selaginella pulvinata (Hook.& Grev.) Maxim. in India. Indian Journal of Forestary 28(4): 389-391.



PUNETHA, N., A.K.PANT & B. S. KHOLIA. 2004. Pteridophytic diversity of Kumaon in ed. T.A.. Sharma and J.S. Dargan Plant diversity of India. 267-280.



B. S. KHOLIA & PUNETHA, N. 2004. The genus Colysis Presl. in western Himalaya. Phytotaxonomny 3:119-123. ISSN No 0972-4206 12 (1),



B. S. KHOLIA & PUNETHA, N. 2002. Diplazium subsinuatum (Wall.ex Hook. et Grev) Tag. In Central Himalaya Phytotatxonomy 2:163-165.



Punetha, N., A. K. Pant & B. S. Kholia. 2003. A report on pteridophyte diversity of Milam (Nanda Devi Biosphere Reserve). In R. R. Das & R. P. Sing (Eds.). Proceedings of the National Seminar on Biodiversity: Conservation and Management. Environmental Planning and Coordination Organization, Bhopal (India). Pp. 143-146.



PUNETHA, N. & B. S. KHOLIA. 1998. Pteridophytic flora of Kumaon: Distribution, Evolution and status. Indian Fern J.15:75 – 79.



B. S. KHOLIA & PUNETHA, N. 1995. Notes on taxonomy and distribution of some Pteridophytes in Kumaon (W.Himalaya) J. Indian Bot. Soc. 74:183 -187.



B. S. KHOLIA & PUNETHA, N. 1993. An enumeration of Athyrium Roth. species from Pithoragarh district of Kumaon (W. Himalaya) in Higher Plants of Indian Subcontinent Add. Ser. of Indian J.For 4:243 – 251.


PUNETHA, N., ABHA SEN & B. S. KHOLIA. 1991. Taxonomic observations on the pteridophytic flora of Pithoragarh district of Kumaon ( N.W. Himalaya):-Fern –allies in B.K. Gupta ed. Higher Plants of Indian Subcontinent (Add. Ser. of Indian J. For.) 3: 125-142.


PUNETHA, N. & B. S. KHOLIA. 1991. Forking of fronds in certain Ferns. J. Bombay nat. Hist. Soc. 88:142-143. ISSN 0006-6982


PUNETHA, N. & B. S. KHOLIA. 1990. Lepisorus pseudoclathratus Ching et Wu, A new record for India. Indian Fern J. 7:40- 42.


PUNETHA, N. & B. S. KHOLIA. 1990. Two new varieties of Christella dentata (Forssk.)Brownsey et Jermy. J.Bombay nat. Hist. Soc. 87:264 – 266.


PUNETHA, N. & B. S. KHOLIA Lepisorus gyriongensis and Lepisorus tibeticus new record for India. Indian Fern J.


PUNETHA, N. & B. S. KHOLIA. 1989. Addition to the Pteridophytic flora of Pithoragarh district of Kumaon (W. Himalaya). NewBotanist. 36: 115-126.


PUNETHA, N. & B. S. KHOLIA. 1989. On the identity and distribution of Oleandra pistillaris (SW) C. Chr. in India. Bull.Bot. Surv. India 31:180 – 181.


PUNETHA, N. & B. S. KHOLIA. 1989. Pnumatopteris nudata (Roxb.) Punetha et Kholia comb novo. J. Bombay nat. Hist. Soc.86:475-479.


PUNETHA, N. & B. S. KHOLIA. 1989. Polystichum makinoi Tag. a new record for India Curr.Sci. 58:317-318.


PUNETHA, N. & B. S. KHOLIA. 1989.A note on cytology of Polystichum luctosum (Kunze) T, Moore Indian Fern J. 5:125-126.


KHULLAR, S. P., PUNETHA, N. & B. S. KHOLIA. 1988. The species of Dryopteris. in the Pithoragarh district of Kumaon (West Himalaya) Indian Fern J.5:127-135.


PUNETHA, N. & B. S. KHOLIA. 1988. Seasonal variations of the fern spores in the air of Pithoragarh town. Him. J. of Env. & Zool. 2:110-113. ISSN: 0970-2903.


PUNETHA, N. & B. S. KHOLIA. 1988. Vittaria himalayensis Ching, A new record for western Himalaya. Curr. Sci 57:346-348.


PUNETHA, N. & B. S. KHOLIA. 1988.On the occurrence of Dryopteris conjugata Ching in the western Himalaya. Indian J.For.11:5340 – 341.


PUNETHA, N. & B. S. KHOLIA. 1987. A list of Polystichum Roth species in the Pithoragarh district of Kumaon (N.W. Himalaya). Bull. Pure & Appl. Sci. 6:3-8.

Dr. B. S. Kholia, Ph. D.
Botanical Survey of India
192- Kaulagarh Road
DEHRADUN- 248 195